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All photos copyright by Martin Holladay and Karyn Patno.

Achilles tang – Acanthurus achilles – Rangiroa
Achilles tang – Acanthurus achilles
Antler coral – Pocillopora grandis – Fakarava
Arc-eye hawkfish – Paracirrhites arcatus
Arc-eye hawkfish – Paracirrhites arcatus – Tikehau
Banded sergeant – Abudefduf septemfasciatus – Fakarava
Bandtail goatfish – Upeneus taeniopterus – Tikehau
Barred thicklip – Hemigymnus fasciatus – Tikehau
Bicolor cleaner wrasse – Labroides bicolor – Rangiroa
Big longnose butterflyfish – Forcipiger longirostris
Bignose unicornfish – Naso vlamingii – Rangiroa
Bignose unicorn fish – Naso vlamingii
Blackbar damsel – Plectroglyphidodon dickii
Blackbar damsel – Plectroglyphidodon dickii – Tikehau
Blacksaddle coral grouper – Plectropomus laevis – Fakarava
Blacksaddle coralgrouper – Plectropomus laevis – Fakarava
Black-saddled toby – Canthigaster valentini – Rangiroa
Blackspot sergeant – Abudefduf sordidus – Rangiroa
Blackspot sergeant – Abudefduf sordidus – Rangiroa
Blacktail snapper – Lutjanus fulvus – Rangiroa
Blacktail snapper – Lutjanus fulvus – Fakarava
Blacktip reef shark – Carcharhinus melanopterus – Fakarava
Blacktip reef shark – Carcharhinus melanopterus
Blacktip reef shark – Carcharhinus melanopterus – Rangiroa
Black triggerfish – Melichthys niger – Tikehau
Bleeker’s parrotfish – Chlorurus bleekeri – Tahiti
Bluebarred parrotfish – Scarus ghobban – Tikehau
Bluefin trevally – Caranx melampygus – Rangiroa
Bluefin trevally – Caranx melampygus – Tikehau
Bluefin trevally – Caranx melampygus
Blue-green chromis – Chromis viridis – Tahiti
Blue-green chromis – Chromis viridis – Tahiti
Bluepatch parrotfish – Scarus forsteni – Tikehau
Bluepatch parrotfish – Scarus forsteni – Tikehau
Bluespine unicornfish – Naso unicornis – Fakarava
Bluespot mullet – Moolgarda seheli – Rangiroa
Bluespot mullet – Moolgarda seheli
Blue-spotted cornetfish – Fistularia commersonii – Fakarava
Bluespotted wrasse – Anampses caeruleopunctatus – Tikehau
Bluespotted wrasse – Anampses caeruleopunctatus – Tikehau
Bluespotted wrasse – Anampses caeruleopunctatus – Tikehau
Bluestreak cleaner wrasse – Labroides dimidiatus – Rangiroa
Bluestripe snapper – Lutjanus kasmira – Fakarava
Bluestripe snapper – Lutjanus kasmira – Fakarava
Blue trevally – Carangoides ferdau – Tikehau
Brassy chub – Kyphosus vaigiensis – Tikehau
Brick soldierfish – Myripristis amaena – Rangiroa
Bridled parrotfish – Scarus frenatus – Tikehau
Bridled parrotfish – Scarus frenatus – Rangiroa
Bridled parrotfish – Scarus frenatus – Tikehau
Brushtail tang – Zebrasoma scopas – Fakarava
Cauliflower coral – Pocillopora meandrina – Fakarava
Checkerboard wrasse – Halichoeres hortulanus – Tikehau
Checkerboard wrasse – Halichoeres hortulanus – Rangiroa
Chevron butterflyfish – Chaetodon trifascialis – Tahiti
Clouded lizardfish – Saurida nebulosa – Fakarava
Clown coris – Coris aygula – Tikehau
Clown coris -Coris aygula – Rangiroa
Clown coris – Coris aygula – Fakarava
Clown coris – juvenile – Coris aygula – Tikehau
Commercial top shell – Rochia nilotica – Rangiroa
Convict surgeonfish – Acanthurus triostegus – Tahiti
Convict surgeonfish – Acanthurus triostegus – Tahiti
Coral grouper – Cephalopholis miniata – Fakarava
Crocodile needlefish – Tylosurus crocodilus – Fakarava
Dash-dot goatfish – Parupeneus barbennus – Rangiroa
Dash-dot goatfish – Parupeneus barbennus – Tikehau
Dash-dot goatfish – Parupeneus barbennus – Rangiroa
Dot-and-dash butterflyfish – Chaetodon pelewensis – Tikehau
Double-spotted queenfish – Scomberoides lysan – Fakarava
Double-spotted queenfish – Scomberoides lysan – Tikehau
Double-spotted queenfish – Scomberoides lysan – Fakarava
Dusky gregory – Stegastes nigricans – Tahiti
Dusky gregory – Stegastes nigricans – Tikehau
Dusky gregory – Stegastes nigricans – Tahiti
Dusky gregory – Stegastes nigricans – Tikehau
Dusky parrotfish – Scarus niger – Rangiroa.jpg
Eclipse butterfllyfish – Chaetodon bennetti – Fakarava
Emperor angelfish – Pomacanthus imperator – Tikehau
Filament-fin parrotfish – Scarus altipinnis – Rangiroa
Filament-fin parrotfish – Scarus altipinnis – Tikehau
Filament-fin parrotfish – Scarus altipinnis – Fakarava
Filament-fin parrotfish – juvenile – Scarus altipinnis – Rangiroa
Fivestripe wrasse – Thalassoma quinquevittatum – Tikehau
Fivestripe wrasse – Thalassoma quinquevittatum – Tikehau
Flagtail grouper – Cephalopholis urodeta
Flagtail grouper – Cephalopholis urodeta – Tikehau
Flagtail triggerfish – Sufflamen chrysopterum – Tikehau
Flame hawkfish – Neocirrhites armatus – Tikehau
Floral wrasse – Cheilinus fasciatus – Rangiroa
Fluted giant clam – Tridacna squamosa – Fakarava
Fourspot butterflyfish – Chaetodon quadrimaculatus
Fourspot butterflyfish – Chaetodon quadrimaculatus – Tikehau
Freckled hawkfish – Paracirrhites forsteri – Fakarava
Giant moray – Gymnothorax javanicus – Rangiroa
Giant moray – Gymnothorax javanicus
Giant moray – Gymnothorax javanicus
Goldsaddle goatfish – Parupeneus cyclostomus – Fakarava
Goldsaddle goatfish – Parupeneus cyclostomus – Fakarava
Great barracuda – Sphyraena barracuda
Great hammerhead shark – Sphyrna mokarran
Great hammerhead shark – Sphyrna mokarran – Tikehau
Great hammerhead shark – Sphyrna mokarran – Tikehau
Guineafowl puffer – Arothron meleagris – Tikehau
Guineafowl puffer – Arothron meleagris – Tikehau
Half-moon triggerfish – Rhinecanthus lunula – Tikehau
Half-moon triggerfish – Rhinecanthus lunula
Halfspotted hawkfish – Paracirrhites hemistictus – Tikehau
Halfspotted hawkfish – Paracirrhites hemistictus – Tikehau
Hawaiian bristletooth – Ctenochaetus hawaiiensis – Tikehau
Hawaiian bristletooth – Ctenochaetus hawaiiensis – Tikehau
Hawksbill sea turtle – Eretmochelys imbricata – Tikehau
Honeycomb grouper – Epinephelus merra – Rangiroa
Humpback snapper – Lutjanus gibbus – Fakarava.jpg
Humpback snapper – Lutjanus gibbus – Rangiroa
Humpback snapper – Lutjanus gibbus – Tikehau
Humpback unicornfish – Naso brachycentron
Humphead wrasse – Cheilinus undulatus – Fakarava
Humphead wrasse – Cheilinus undulatus
Humpnose bigeye bream – Monotaxis grandoculis – Fakarava
Humpnose bigeye bream – Monotaxis grandoculis – Rangiroa
Humpnose bigeye bream – Monotaxis grandoculis – Tikehau
Humpnose bigeye bream – Monotaxis grandoculis – Fakarava
Indo-Pacific Christmas tree worm – Spirobranchus corniculatus – Fakarava
Indo-Pacific sergeant – Abudefduf vaigiensis – Rangiroa
Island goatfish – Parupeneus insularis – Tikehau
Johnston damsel – Plectroglyphidodon johnstonianus
Keeltail needlefish – Platybelone argalus – Tikehau
Ladder wrasse – Thalassoma trilobatum – Tikehau
Lagoon triggerfish – Rhinecanthus aculeatus – Tahiti
Lagoon triggerfish – Rhinecanthus aculeatus – Rangiroa
Lemonpeel angelfish – Centropyge flavissima
Leopard coral grouper – Plectropomus leopardus
Leopard sea cucumber – Bohadschia argus – Tahiti
Leopard sea cucumber – Bohadschia argus – Tahiti.jpg
Longface emperor – Lethrinus olivaceus – Tikehau
Longnose tang – Zebrasoma rostratum – Tikehau
Longnose tang – Zebrasoma rostratum – Tikehau
Mackerel scad – Decapterus macarellus – Tikehau
Manybar goatfish – Parupeneus multifasciatus – Rangiroa
Many-bar goatfish – Parupeneus multifasciatus – Rangiroa
Manybar goatfish – Parupeneus multifasciatus – Rangiroa
Manybar goatfish – Parupeneus multifasciatus -Fakarava
Masked bannerfish – Heniochus monoceros – Rangiroa
Milkfish – Chanos chanos – Tikehau
Mimic surgeonfish – Acanthurus pyroferus – Rangiroa
Mimic surgeonfish – Acanthurus pyroferus – Fakarava
Mimic surgeonfish – Acanthurus pyroferus – Fakarava
Moorish idol – Zanclus cornutus- Tahiti
Moorish idol – Zanclus cornutus – Rangiroa
Mushroom coral – Fungia fungites – Rangiroa
Ocellated eagle ray – Aetobatus ocellatus – Rangiroa
Onespot snapper – Lutjanus monostigma – Tikehau
Onespot snapper – Lutjanus monostigma – Fakarava
Orangeband surgeonfish – Acanthurus olivaceus – Rangiroa
Orangeband suregeonfish – subadult – Acanthurus olivaceus – Fakarava
Orangeband surgeonfish – Acanthurus olivaceus – Fakarava
Orange-lined triggerfish – Balistapus undulatus – Tikehau
Orangespine unicornfish – Naso elegans
Ornate butterflyfish – Chaetodon ornatissimus – Tahiti
Pacific bird wrasse – Gomphosus varius – Tikehau
Pacific bird wrasse – Gomphosus varius – Tikehau
Pacific bullethead parrotfish – Chlorurus spilurus- Fakarava
Pacific bullethead parrotfish – Chlorurus spilurus – Rangiroa
Pacific bullethead parrotfish – Chlorurus spilurus – Fakarava
Pacific bullethead parrotfish – Chlorurus spilurus – Fakarava
Pacific bullethead parrotfish – Chlorurus spilurus – Fakarava
Pacific bullethead parrotfish – Chlorurus spilurus- Fakarava
Pacific bullethead parrotfish – Chlorurus spilurus – Fakarava
Pacific double-saddle butterflyfish – Chaetodon ulietensis – Fakarava
Pacific longnose parrotfish – Hipposcarus longiceps – Tikehau
Pacific sailfin tang – Zebrasoma veliferum – Rangiroa
Pacific peacock flounder – Bothus mancus – Tikehau
Pacific surge damsel – Chrysiptera leucopoma – Rangiroa
Palenose parrotfish – Scarus psittacus – Rangiroa
Palenose parrotfish – Scarus psittacus – Tikehau
Pale-tailed bristletooth – Ctenochaetus flavicauda – Rangiroa
Paletail unicorn fish – Naso brevirostris – Rangiroa
Paletail unicorn fish – Naso brevirostris – Rangiroa
Peacock damsel -Pomacentrus pavo – Fakarava
Peacock grouper – Cephalopholis argus – Tikehau
Pennant bannerfish – Heniochus chrysostomus – Fakarava
Pennant bannerfish – Heniochus chrysostomus – Fakarava
Pin-cushion sea star – Culcita novaeguineae – Fakarava
Pineapple sea cucumber – Thelenota ananas – Rangiroa
Pinktail triggerfish – Melichthys vidua – Fakarava
Pinktail triggerfish – Melichthys vidua – Tikehau
Polynesian humbug dascyllus – Dascyllus emamo – Fakarava
Polynesian humbug dascyllus – Dascyllus emamo – Fakarava
Polynesian humbug dascyllus – Dascyllus emamo – Fakarava
Porcupine fish – Diodon hystrix – Tikehau
Raccoon butterflyfish – Chaetodon lunula – Tikehau
Raccoon-face butterflyfish – Chaetodon lunula – Rangiroa
Raccoon-face butterflyfish – Chaetodon lunula – Rangiroa
Redfin bream – Monotaxis heterodon – Fakarava
Redfin butterflyfish – Chaetodon lunulatus – Fakarava
Redfin butterflyfish – Chaetodon lunulatus – Fakarava.jpg
Redfin bream – Monotaxis heterodon – Rangiroa
Redfin bream – Monotaxis heterodon – Fakarava
Redlip parrotfish – Scarus rubroviolaceus
Redshoulder wrasse – Stethojulis bandanensis – Tikehau
Redshoulder wrasse – immature – Fakarava
Red snapper – Lutjanus bohar – Tikehau
Redtooth triggerfish – Odonus niger – Tikehau
Reefcrest parrotfish – Chlorurus frontalis – Rangiroa
Reefcrest parrotfish – Chlorurus frontalis – Rangiroa
Reefcrest parrotfish – Chlorurus frontalis – Rangiroa
Reefcrest parrotfish – Chlorurus frontalis – Tikehau
Reefcrest parrotfish – Chlorurus frontalis – Tikehau
Reefcrest parrotfish – Chlorurus frontalis – Tikehau
Reefcrest parrotfish – Chlorurus frontalis – Tikehau
Reefcrest parrotfish – Chlorurus frontalis – Tikehau.jpg
Reefcrest parrotfish school – Chlorurus frontalis – Tikehau
Regal angelfish – Pygoplites diacanthus – Rangiroa
Reticulated butterflyfish – Chaetodon reticulatus – Rangiroa
Reticulated butterflyfish – Chaetodon reticulatus – Rangiroa
Ringtail wrasse – Oxycheilinus unifasciatus – Rangiroa
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Rockmover wrasse – Novaculichthys taeniourus – Tikehau
Rockmover wrasse – Novaculichthys taeniourus – Rangiroa
Saber squirrelfish – Sargocentron spiniferum – Fakarava
Saddled butterflyfish – Chaetodon ephippium – Tikehau
Scissortail sergeant – Abudefduf sexfasciatus – Fakarava
Scythe triggerfish – Sufflamen bursa – Tikehau
Sharksucker – Echeneis naucrates – Fakarava
Sharksucker – Echeneis naucrates – Fakarava
Sixbar wrasse – Thalassoma hardwicke – Tahiti
Slingjaw wrasse – Epibulus insidiator – Fakarava
Slingjaw wrasse – Epibulus insidiator – Tahiti
Slingjaw wrasse – Epibulus insidiator – Rangiroa
Small-spotted dart – Trachinotus baillonii – Tikehau
Solander’s toby – Canthigaster solandri – Rangiroa
Snowflake moray – Echidna nebulosa – Tikehau.jpg
Speckled butterflyfish – Chaetodon citrinellus – Tikehau
Speckled butterflyfish – Chaetodon citrinellus – Tahiti
Spotfin squirrelfish – Neoniphon sammara – Rangiroa
Spotted parrotfish – Cetoscarus ocellatus – Rangiroa
Spotted parrotfish – Cetoscarus ocellatus – Rangiroa
Spotted parrotfish – Cetoscarus ocellatus – Fakarava
Spotted parrotfish – Cetoscarus ocellatus – Rangiroa
Spotted parrotfish juvenile – Cetoscarus ocellatus – Fakarava
Squaretail mullet – Ellochelon vaigiensis – Fakarava
Squaretail mullet – Ellochelon vaigiensis – Rangiroa
Stareye parrotfish – Calotomus carolinus – Tikehau
Star puffer – Arothron stellatus – Tikehau
Steephead parrotfish – Chlorurus microrhinos – Fakarava
Steephead parrotfish – Chlorurus microrhinos – Tikehau
Steephead parrotfish – Chlorurus microrhinos – Fakarava
Steephead parrotfish – Chlorurus microrhinos – Fakarava
Steephead parrotfish red variation – Chlorurus microrhinos – Rangiroa
Steephead parrotfish red variation – Chlorurus microrhinos – Rangiroa
Striped surgeonfish – Acanthurus lineatus – Tikehau
Surge wrasse – Thalassoma purpureum – Tikehau
Surge wrasse immature – Thalassoma purpureum – Tikehau
Surge wrasse – Thalassoma purpureum – Rangiroa
Swarthy parrotfish – Scarus niger – Rangiroa
Tahitian butterflyfish – Chaetodon trichrous – Rangiroa
Tahitian butterflyfish – Chaetodon trichrous – Rangiroa
Tahitian stingray – Pateobatis fai – Tikehau
Teardrop butterfly fish – Chaetodon unimaculatus – Fakarava
Threadfin butterflyfish – Chaetodon auriga – Fakarava
Three-spot dascyllus juvenile – Dascyllus trimaculatus – Tahiti
Threespot wrasse – Halichoeres trimaculatus – Fakarava
Threespot wrasse – Halichoeres trimaculatus – Fakarava
Titan triggerfish – Balistoides viridencens – Tikehau
Titan triggerfish – Balistoides viridencens – Fakarava
Titan triggerfish – Balistoides viridencens – Rangiroa
Threadfin butterflyfish – Chaetodon auriga – Fakarava
Tripletail wrasse – Cheilinus trilobatus – Rangiroa
Tripletail wrasse – Cheilinus trilobatus – Rangiroa
Tripletail wrasse – Cheilinus trilobatus – Tikehau
Tripletail wrasse – Cheilinus trilobatus – Tahiti
Two-spined angelfish – Centropyge bispinosa – Rangiroa
Two-spot red snapper – Lutjanus bohar – Tikehau
Wedgetail triggerfish – Rhinecanthus rectangulus – Tikehau
Weedy surge wrasse – Halichoeres margaritaceus – Tikehau
Whitebar gregory – Stegastes albifasciatus – Tikehau
Whitebelly toby – Canthigaster bennetti – Fakarava
Whitecheek surgeonfish – Acanthurus nigricans – Fakarava
Whitecheek surgeonfish – Acanthurus nigricans – Fakarava
Whitecheek surgeonfish – Acanthurus nigricans – Fakarava
Whitemouth moray – Gymnothorax meleagris – Tahiti
White-spotted hawkfish – Cirrhitus pinnulatus – Tikehau
White-spotted puffer – Arothron hispidus – Tahiti
White-spotted surgeonfish – Acanthurus guttatus – Tikehau
Whitetail surgeonfish – Acanthurus thompsoni – Fakarava
Whitetip reef shark – Triaenodon obesus – Fakarava
Yellowband parrotfish – Scarus schlegeli – Rangiroa
Yellowband parrotfish – Scarus schlegeli – Rangiroa
Yellowband parrotfish – Scarus schlegeli – Rangiroa
Yellow boxfish – Ostracion cubicus – Tahiti
Yellow boxfish – Ostracion cubicus – Fakarava
Yellow-edged lyretail -Variola louti – Tikehau
Yellow-edged lyretail -Variola louti – Tikehau
Yellowfin goatfish -Mulloidichthys vanicolensis – Tahiti
Yellowfin goatfish – Mulloidichthys vanicolensis – Fakarava
Yellowmargin triggerfish – Pseudobalistes flavimarginatus – Rangiroa
Yellowmargin triggerfish – Pseudobalistes flavimarginatus – Rangiroa
Yellowstripe goatfish – Mulloidichthys flavolineatus – Rangiroa
Yellowstripe goatfish – Mulloidichthys flavolineatus – Fakarava
Yellowstripe goatfish – Mulloidichthys flavolineatus – Fakarava
Yellowstripe goatfish – Mulloidichthys flavolineatus – Tikehau
Yellowtail coris – Coris gaimard – Tikehau