Maldives

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All photos copyright by Martin Holladay and Karyn Patno.

Acropora digitifera coral – Acropora digitifera
Andaman butterflyfish – Chaetodon adamanensis
Axilspot hogfish – Bodianus axillaris
Axilspot hogfish – Bodianus axillaris
Banded coral shrimp – Stenopus hispidus
Banded sergeant – Abudefduf septemfasciatus
Barred goatfish – Parupeneus trifasciatus
Barred thicklip wrasse – Hemigymnus fasciatus
Barred thicklip wrasse – Hemigymnus fasciatus
Bicolor blenny – Ecsenius bicolor
Bicolor cleaner wrasse – Labroides bicolor
Big longnose butterflyfish – Forcipiger longirostris
Bignose unicornfish – Vlaming’s unicornfish – Naso vlamingii
Bignose unicornfish – Vlaming’s unicornfish – Naso vlamingii
Bignose unicornfish – Vlaming’s unicornfish – Naso vlamingii
Bird wrasse – Gomphosus caeruleus
Bird wrasse – Gomphosus caeruleus
Black-and-white snapper – Macolor niger
Black-backed butterflyfish – Blackback butterflyfish – Chaetodon melannotus
Black-blotched porcupinefish – Diodon liturosus
Blackeye rabbitfish – Siganus puelloides
Blackeye thicklip wrasse – Hemigymnus melapterus
Blackfin grouper – Cephalopholis nigripinnis
Blackfin squirrelfish – Neoniphon opercularis
Black pyramid butterflyfish – Hemitaurichthys zoster
Black-saddle coral grouper – Plectropomus laevis
Blacksaddle toby -Saddled sharpnose puffer – Canthigaster valentini
Black-spotted pufferfish – Arothron nigropunctatus
Blackspotted sea cucumber – Pearsonothuria graeffei
Blackspotted sea cucumber – Pearsonothuria graeffei
Blacktip reef shark – Carcharhinus melanopterus
Blacktip reef shark – Carcharhinus melanopterus
Blackwedged butterflyfish – Chaetodon falcula
Blue and yellow fusilier – Caesio teres
Blue blanquillo – Malacanthus latovittatus
Blue blanquillo – Malacanthus latovittatus
Blue-faced angelfish – Pomacanthus xanthometopon
Bluefin trevally – Caranx melampygus
Blue-green chromis – Chromis viridis
Bluelined wrasse – Stethojulis albovittata
Bluelined wrasse – Stethojulis albovittata
Bluespine unicornfish – Naso unicornis
Bluespotted cornetfish – Fistularia commersonii
Bluestreak cleaner wrasse – Labroides dimidiatus
Bluestreak cleaner wrasse – Labroides dimidiatus
Bluestreak cleaner wrasse – Labroides dimidiatus
Blue-striped snapper – Lutjanus kasmira
Blunthead wrasse – Thalassoma amblycephalum
Bridled monocle bream – Two-lined monocle bream – Scolopsis bilineata.jpg
Bridled parrotfish – Scarus frenatus
Bridled parrotfish – Scarus frenatus
Bridled parrotfish – Scarus frenatus
Bridled parrotfish juvenile – Scarus frenatus
Bridled parrotfish – Scarus frenatus
Brushtail tang – Zebrasoma scopas
Camouflage grouper – Epinephelus polyphekadion
Checkerboard wrasse – Halichoeres hortulanus
Checkerboard wrasse – Halichoeres hortulanus
Checkerboard wrasse – Halichoeres hortulanus
Chevron butterflyfish – Chaetodon trifascialis
Chiragra conch – Harpago chiragra
Circular batfish – Platax orbicularis
Clark’s anemonefish – Amphiprion clarkii
Clown coris – Coris aygula
Clown triggerfish – Balistoides conspriculum
Collared butterflyfish – Chaetodon collare
Common mushroom coral – Fungia fungites
Convict surgeonfish – Acanthurus triostegus
Coral grouper – Cephalopholis miniata
Coral grouper – Vermillion rock cod – Cephalopholis miniata
Crescent-tail bigeye – Priacanthus hamrur
Crescent-tail bigeye – Priacanthus hamrur
Crimson solderifish – Myripristis murdjan
Dash-dot goatfish – Parupeneus barberinus
Day octopus – Octopus cyanea
Dogtooth tuna – Gymnosarda unicolor
Double-spotted queenfish – Scomberoides lysan
Dusky angelfish – Multispine angelfish – Centropyge multispinis
Dusky gregory – Stegastes nigricans
Dusky gregory – Stegastes nigricans
Dusky gregory – Stegastes nigricans
Dusky parrotfish – Scarus niger
Dusky parrotfish – Scarus niger
Dusky wrasse – Halichoeres marginatus
Eclipse butterflyfish – Chaetodon bennetti
Elegant unicornfish – Naso elegans
Emperor angelfish – Imperial angelfish – Pomacanthus imperator
Emperor angelfish – Imperial angelfish – Pomacanthus imperator
Eyestripe surgeonfish – Acanthurus nigricauda
Festive parrotfish – Scarus festivus
Flagtail triggerfish – Sufflamen chrysopterum
Fluted giant clam – Tridacna squamosa
Fluted giant clam – Tridacna squamosa
Fluted giant clam – Tridacna squamosa
Forktail rabbitfish – Siganus argenteus
Forktail rabbitfish – Siganus argenteus
Four-saddled grouper – Epinephelus spilotoceps
Freckled hawkfish – Paracirrhites forsteri
Fringelip mullet – Crenimugil crenilabis
Galathea sea star – Brown mesh sea star – Nardoa galatheae
Giant moray – Gymnothorax javanicus
Giant trevally – Caranx ignobilis
Golden chromis – Ternate chromis – Chromis ternatensis
Golden trevally – Gnathanodon speciosus
Goldring bristletooth surgeonfish – Ctenochaetus truncatus
Gold-saddle goatfish – Yellow-saddle goatfish – Parupeneus cyclostomus
Gold-saddle goatfish – Yellow-saddle goatfish – Parupeneus cyclostomus
Gold-striped emperor – Gnathodentex aureolineatus
Granulated sea star – Choriaster granulatus
Great barracuda – Sphyraena barracuda
Green barrel sea squirt – Didemnum molle
Greencheek parrotfish – Scarus prasiognathus
Greenfish sea cucumber – Stichopus chloronotus
Green jobfish – Aprion virescens
Green sea turtle – Chelonia mydas
Guilding’s sea star – Linckia guildingi
Guineafowl puffer – Arothron meleagris
Halfmoon triggerfish – Rhinecanthus lunula
Hawksbill sea turtle – Eretmochelys imbricata
Honeycomb grouper – Epinephelus merra
Humbug dascyllus – Dascyllus aruanus
Humpback snapper – Lutjanus gibbus
Humpback unicornfish – Naso brachycentron
Humphead wrasse – Napolean wrasse – Cheilinus undulatus
Humpnose bigeye bream – Monotaxis grandoculis
Humpnose bigeye bream – Monotaxis grandoculis
Imposter fangblenny – Imposter sabretooth blenny – Plagiotremus phenax
Indian bannerfish – Heniochus pleurotaenia
Indian bullethead parrotfish – Chlorurus sordidus
Indian damselfish – Pomacentrus indicus
Indian damselfish – Pomacentrus indicus
Indian dascyllus – Dascyllus carneus
Indian half-and-half chromis – Pycnochromis fieldi
Indian longnose parrotfish – Hipposcarus harid
Indian longnose parrotfish – Hipposcarus harid
Indian longnose parrotfish – Hipposcarus harid
Indian longnose parrotfish – Hipposcarus harid
Indian Ocean pinstriped wrasse – Halichoeres vrolikii
Indian parrotfish – Pink-margined parrotfish – Chlorurus capistratoides
Indian sailfin tang – Zebrasoma desjardinii
Indian sea star – Fromia indica
Indian triggerfish – Melichthys indicus
Indo-Pacific sergeant – Abudefduf vaigiensis
Jewel damselfish – Plectroglyphidodon lacrymatus
Johnston damselfish – Plectroglyphidodon johnstonianus
Kawakawa – Mackerel tuna – Euthynnus affinis
Kleins butterflyfish – Blacklip butterflyfish – Chaetodon kleinii
Lagoon triggerfish – Rhinecanthus aculeatus
Leopard grouper – Leopard rock cod – Cephalopholis leopardus
Leopard grouper – Cephalopholis leopardus
Lieutenant surgeonfish – Acanthurus tennenti
Lined bristletooth – Ctenochaetus striatus
Lined butterflyfish – Chaetodon lineolatus
Little combtooth blenny – Ecsenius minutus
Longbarbel goatfish – Parupeneus macronemus
Longnose butterflyfish – Forcipiger flavissimus
Longnose butterflyfish – Forcipiger flavissimus
Longnose butterflyfish – Forcipiger flavissimus
Longtail silverbiddy – Gerres longirostris
Lunar fusilier – Caesio lunaris
Lunar-tailed grouper – Yellow-edged lyretail – Variola louti
Lyretail anthias – Pseudanthias squamipinnis
Madagascar butterflyfish – Seychelles butteflyfish – Chaetodon madagaskariensis
Maldives triplefin – Helcogramma maldivensis
Maldivian anemonefish – Blackfin anemonefish – Amphirpion nigripes
Maldivian anemonefish – Blackfin anemonefish – Amphirpion nigripes
Masked bannerfish – Heniochus monoceros
Meyers butterflyfish – Chaetodon meyeri
Midnight snapper – Macolor macularis
Midnight snapper juvenile – Macolor macularis
Moon wrasse – Crescent wrasse – Thalassoma lunare
Moorish idol – Zanclus cornutus
Neon damselfish – Pomacentrus caeruleus
Neon fusilier – Pterocaesio tile
Noduled sea star – Fromia nodosa
Ocellated eagle ray – Aetobatus ocellatus
Onespot snapper – Lutjanus monostigma
Orange anthias – Pseudanthias squamipinnis
Orange anthias – Pseudanthias squamipinnis
Orange cup coral – Tubastraea coccinea
Orange-fin emperor – Lethrinus erythracanthus
Orange-lined triggerfish – Balistapus undulatus
Orangespine unicornfish – Elegant unicornfish – Naso elegans
Orange-stripe emperor – Lethrinus obsoletus
Oriental sweetlips – Plectorhinchus vittatus
Ornate emperor – Lethrinus ornatus
Oval butterflyfish -Indian redfin butterflyfish – Chaetodon trifasciatus
Pacific peacock flounder – Bothus mancus
Painted spiny lobster – Panulirus versicolor
Paletail unicornfish – Naso brevirostris
Paletail unicornfish – Naso brevirostris
Peacock damselfish – Pomacentrus pavo
Peacock grouper – Cephalopholis argus
Powderblue surgeonfish – Acanthurus leucosternon
Queen coris juvenile – Coris formosa
Raccoon butterflyfish – Chaetodon lunula
Redbreasted Maori wrasse – Cheilinus fasciatus
Redlip parrotfish – Scarus rubroviolaceus
Redmouth grouper – Aethaloperca rogaa
Redshoulder wrasse – Stethojulis bandanensis
Red snapper – Lutjanus bohar
Red snapper – Lutjanus bohar
Redtooth triggerfish – Red-toothed triggerfish – Odonus niger
Reef lizardfish – Variegated lizardfish – Synodus variegatus
Reef needlefish – Strongylura incisa
Regal angelfish – Pygoplites diacanthus
Robust sea squirt – Atriolum robustum
Rockmover wrasse – Novaculichthys taeniourus
Roundhead parrotfish – Chlorurus strongylocephalus
Roundhead parrotfish – Chlorurus strongylocephalus
Roundhead parrotfish – Chlorurus strongylocephalus
Ruddy fusilier – Pterocaesio pisang
Saber squirrelfish – Sargocentron spiniferum
Saddleback butterflyfish – Chaetodon falcula
Scythe triggerfish – Sufflamen bursa
Sebree’s micro-goby – Striped dwarf goby – Eviota sebreei
Shadowfin soldierfish – Myripristis adusta
Sharksucker – Echeneis naucrates
Silty cardinalfish – Ostorhinchus properuptus
Silverspot squirrelfish – Sargocentron caudimaculatum
Silverspot squirrelfish – Sargocentron caudimaculatum
Singlespine unicornfish – Onespine unicornfish – Naso thynnoides
Sixbar wrasse – Thalassoma hardwicke
Sixline soapfish – Grammistes sexlineatus
Sixspot goby – Valenciennea sexguttata
Sixspot rock cod – Saddle grouper – Cephalopholis sexmaculata
Sixstripe wrasse – Sixline wraasse – Pseudocheilinus hexataenia
Slate pencil urchin – Heterocentrotus mamillatus
Sleek unicornfish – Naso hexacanthus
Sleek unicornfish – Naso hexacanthus
Slender grouper – Anyperodon leucogrammicus
Slender silverbiddy – Gerres oblongus
Slingjaw wrasse – Epibulus insidiator
Slingjaw wrasse – Epibulus insidiator
Smallspotted dart – Trachinotus bailloni
Smalltooth jobfish – Aphareus furca
Smith’s fangblenny – Meiacanthus smithi
Snowflake moray – Clouded moray – Echidna nebulosa
Speckled butterflyfish – Chaetodon citrinellus
Spiny cushion sea star – Culcita schmideliana
Spiny cushion sea star – Culcita schmideliana
Spotfin lionfish – Pterois antennata
Spotfin squirrelfish – Neoniphon sammara
Spotnape butterflyfish – Chaetodon oxycephalus
Spot-tail sand perch – Spotted sandperch – Parapercis hexophtalma
Spotted boxfish – male – Ostracion meleagris
Spotted coral grouper – Plectropomus maculatus
Spotted Linckia sea star – Linckia multiflora
Spotted Linckia sea star – Linckia multiflora
Spotted Linckia sea star – Linckia multiflora
Spotted Linckia sea star – Linckia multiflora
Spotted parrotfish – Cetoscarus ocellatus
Spotted parrotfish – Cetoscarus ocellatus
Spotted parrotfish – Cetoscarus ocellatus
Spotted porcupinefish – Diodon hystrix
Spotted porcupinefish – Diodon hystrix
Spotted sandperch – Parapercis millepunctata
Square-tail coral grouper – Plectropomus areolatus
Square-tail coral grouper – Plectropomus areolatus
Starry rabbitfish – Siganus stellatus
Schooling bannerfish – Heniochus diphreutes
Striped surgeonfish – Lined surgeonfish – Acanthurus lineatus
Surge damselfish – Chrysiptera brownriggii
Thousand-pores sea star – Fromia milleporella
Threadfin butterflyfish – Chaetodon auriga
Threespot angelfish – Apolemichthys trimaculatus
Threespot angelfish – Apolemichthys trimaculatus
Thumbprint emperor – Lethrinus harak
Titan triggerfish – Balistoides viridescens
Tongue coral – Herpolitha limax
Topsail chub – Highfin chub – Kyphosus cinerascens
Topsail chub – Highfin chub – Kyphosus cinerascens
Triangular butterflyfish – Chaetodon triangulum
Triangular butterflyfish – Chaetodon triangulum
Tricolor parrotfish – Scarus tricolor
Tripletail wrasse – Cheilinus trilobatus
Tripletail wrasse – Cheilinus trilobatus
Twin demoiselle – Neopomacentrus sororius
Twospot damselfish – Twospot demoiselle – Chrysiptera biocellata
Twospot red snapper – Lutjanus bohar
Two-tone dartfish – Ptereleotris evides
Vagabond butterflyfish – Chaetodon vagabundus
Variable bushy feather star – Comaster schlegelii
Variable bushy feather star – Comaster schlegelii
Violet soldierfish – Myripristis violacea
Violet soldierfish – Myripristis violacea
Vrolik’s wrasse – Indian Ocean pinstriped wrasse – Halichoeres vrolikii
White-breasted sergeant – Green damselfish – Amblyglyphidodon indicus
Whiteline lionfish – Clearfin lionfish – Pterois radiata
Whitespotted sharpnose puffer – Honeycomb toby – Canthigaster janthinoptera
Whitetail damselfish – Pomacentrus chrysurus
Whitetip reef shark – Triaenodon obesus
Wirenet filefish – Cantherhines pardalis
Yellowback anthias – Mirolabrichthys evansi
Yellow-edged lyretail – Variola louti
Yellowhead butterflyfish – Chaetodon xanthocephalus
Yellowmask surgeonfish – Elongate surgeonfish – Acanthurus mata
Yellowstripe goatfish – Mulloidichthys flavolineatus
Yellowstripe goatfish – Mulloidichthys flavolineatus
Yellowstriped cardinalfish – Ostorhinchus cyanosoma
Yellowtail sergeant – Abudefduf notatus
Zigzag wrasse – Halichoeres scapularis
Zigzag wrasse – Halichoeres scapularis