Guam and Palau

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All photos copyright by Martin Holladay and Karyn Patno.

Acropora gemmifera coral – Acropora gemmifera – Guam
Anchor tuskfish – Choerodon anchorago – Palau
Andaman butterflyfish – Chaetodon andamanensis – Palau
Arc-eye hawksfish – Paracirrhites arcatus – Guam
Arc-eye hawkfish – Paracirrhites arcatus – Palau
Banded goby – Amblygobius phalaena – Guam
Banded goby – Amblygobius phalaena – Palau
Banded goby – Amblygobius phalaena – Palau
Banded goby – Amblygobius phalaena – Palau
Banded sergeant – Abudefduf septemfasciatus – Guam
Banded shrimpgoby – Cryptocentrus cinctus – Palau
Banded shrimpgoby – Cryptocentrus cinctus – Palau
Barred blenny – Cirripectes polyzona – Guam
Barred blenny – Cirripectes polyzona – Guam
Barred rabbitfish – Siganus doliatus – Palau
Barred thicklip – Hemigymnus fasciatus – Guam
Bicolor angelfish – Centropyge bicolor – Palau
Bicolor chromis – Chromis margaritifer – Palau
Bicolor fangblenny – Plagiotremus laudandus – Palau
Bigfin reef squid – Sepioteuthis lessoniana – Palau
Bignose unicornfish – Naso vlamingii – Guam
Bignose unicornfish – – Naso vlamingii – Palau
Black coralgoby – Paragobiodon melanosomus – Palau
Black damsel – Neoglyphidodon melas – Palau
Black triggerfish – Melichthys niger – Guam
Black-axil chromis – Chromis antripectoralis – Guam
Black-axil chromis – Chromis antripectoralis – Palau
Black-backed butterflyfish – Chaetodon melannotus – Guam
Black-backed butterflyfish – Chaetodon melannotus – Palau
Black-banded snapper – Lutjanus semicinctus – Palau
Blackbar damsel – Plectroglyphidodon dickii – Guam
Blackbarred halfbeak – Hemiramphus far – Guam
Blackbelt hogfish – Bodianus mesothorax – Palau
Black damsel – Neoglyphidodon melas – Palau
Black damsel – Neoglyphidodon melas – Palau
Blackeye thicklip – Hemigymnus melapterus – Guam
Blackeye thicklip – Hemigymnus melapterus – Palau
Blackeye thicklip – Hemigymnus melapterus – Palau
Blacklip butterflyfish – Chaetodon kleinii – Palau
Blackpatch triggerfish – Rhinecanthus verrucosus – Palau
Blacksaddle coral grouper – Plectropomus laevis – Palau
Blacksaddle grouper – Epinephelus howlandi – Palau
Black-saddled toby – Canthigaster valentini – Palau
Blackspot cleaner wrasse – Labroides pectoralis – Palau
Blackspot sergeant – Abudefduf sordidus – Guam
Blackspot snappe – Ehrenberg’s snapper – Lutjanus ehrenbergii – Palau
Blackspotted puffer – Arothron nigropunctatus – Guam
Blackspotted puffer – Arothron nigropunctatus – Palau
Blackspotted sea cucumber – Pearsonothuria graeffei – Palau
Blackstreak surgeonfish – Eyeline surgeonfish – Acanthurus nigricauda – Palau
Black-tail sergeant – Abudefduf lorenz – Palau
Banded sergeant – Abudefduf septemfasciatus – Guam
Blacktail snapper – Lutjanus fulvus – Guam
Blacktail snapper – Lutjanus fulvus – Palau
Blacktip reef shark – Carcharhinus melanopterus – Guam
Blacktip reef shark – Carcharhinus melanopterus – Palau
Blacktip reef shark – Carcharhinus melanopterus – Palau
Blackvent damsel – Dischistodus melanotus -Palau
Bleekers parrotfish initial phase – Chlorurus bleekeri – Palau
Bleeker’s parrotfish – Chlorurus bleekeri – Palau
Blotcheye soldierfish – Myripristis kuntee – Guam
Blue-and-yellow fusilier – Caesio teres – Palau
Blueback damselfish – Pomacentrus simsiang – Palau
Blue damsel – Pomacentrus pavo – Palau
Blue damsel – Pomacentrus pavo – Palau
Blue damsel – Pomacentrus pavo – Palau
Blue devil – Chrysiptera cyanea – Palau
Bluefin trevally – Caranx melampygus – Guam
Bluefin trevally – Caranx melampygus – Palau
Blue-green chromis – Chromis viridis – Guam
Blue-green chromis – Chromis viridis – Palau
Blue sea star – Linckia laevigata – Guam
Blue sea star – Linckia laevigata – Palau
Bluespine unicornfish – Naso unicornis – Guam
Bluespine unicornfish – Naso unicornis – Palau
Bluespine unicornfish – Naso unicornis – Palau
Bluespot damselfish – Chrysiptera oxycephala – Palau
Bluespot mullet – Moolgarda seheli – Palau
Blue-spotted wrasse – Anampses caerulepunctatus – Guam
Bluestreak cleaner wrasse – Labroides dimidiatus – Guam
Bluestreak cleaner wrasse – Labroides dimidiatus – Palau
Blue streak fusilier – Neon fusilier – Pterocaesio tile – Palau
Bluestreak goby – Valenciennea strigata – Guam
Blue trevally – Barred trevally – Carangoides ferdau – Palau
Boring giant clam – Tridacna crocea – Guam
Brassy chub – Brassy drummer – Kyphosus vaigiensis – Palau
Brassy trevally – Caranx papuensis – Guam
Brassy trevally – Caranx papuensis – Palau
Brick soldierfish – Myripristis amaena – Palau
Bridled monocle bream juvenile – Scolopsis bilineata – Palau
Bridled monocle bream juvenile – Scolopsis bilineata – Palau
Bridled parrotfish (intermediate phase) – Scarus frenatus – Palau
Bridled parrotfish (intermediate phase) – Scarus frenatus – Palau
Broadclub cuttlefish – Ascarosepion latimanus – Palau
Broad cowtail stingray – Pastinachus ater – Palau
Broom filefish – Amanses scopas – Palau
Brown curryfish sea cucumber – Zebra sea cucumber – Stichopus vastus – Palau
Brown-marbled grouper – Flowery rockcod – Epinephelus fuscoguttatus – Palau
Brown surgeonfish – Dusky surgeonfish – Acanthurus nigrofuscus – Guam
Brushtail tang – Zebrasoma scopas – Palau
Bubble coral – Plerogyra sinuosa – Palau
Bumphead parrotfish – Bolbometopon muricatum – Palau
Bumphead parrotfish – Bolbometopon muricatum – Palau
Bumphead parrotfish (initial phase) – Bolbometopon muricatum – Palau
Camouflage grouper – Epinephelus polyphekadion – Palau
Carnation coral – Dendronephthya sp. – Palau
Chain-lined wrasse – Halichoeres leucurus – Palau
Chain-lined wrasse – Halichoeres leucurus – Palau
Checkerboard wrasse – Halichoeres hortulanus – Guam
Checkerboard wrasse – Halichoeres hortulanus – Palau
Checkered snapper – Lutjanus decussatus – Palau
Cheek-lined wrasse – Oxycheilinus diagrammus – Guam
Chevroned butterflyfish – Chaetodon trifacialis – Guam
Chevroned butterflyfish – Chaetodon trifacialis – Palau
Chocolate chip sea star – Protoreaster nodosus – Palau
Chocolate grouper – Cephalopholis boenak – Palau
Cigar wrasse – Cheilio inermis – Palau
Circular spadefish – Circular batfish – Platax orbicularis – Palau
Clown triggerfish – Balistoides conspricillum – Palau
Common spider conch – Lambis lambis – Palau
Common spider conch – Lambis lambis – Palau
Convict surgeonfish – Acanthurus triostegus – Palau
Convict surgeonfish – Acanthurus triostegus – Palau
Convict surgeonfish – Acanthurus triostegus – Guam
Coral grouper – Cephalopholis miniata – Palau
Coral rabbitfish – Siganus corallinus – Palau
Cornetfish – Bluespotted cornetfish – Fistularia commersonii – Palau
Crescent wrasse – Moon wrasse – Thalassoma lunare – Palau
Crimson soldierfish – Myripristis murdjan – Palau
Darkcap parrotfish – Scarus oviceps – Palau
Dash-dot goatfish – Parupeneus barbennus – Guam
Dash-dot goatfish – Parupeneus barberinus – Palau
Diagonal-banded sweetlips – Plectorhinchus lineatus – Palau
Dotted butterflyfish – Chaetodon semion – Palau
Double-lined mackerel – Grammatorcynus bilineatus – Palau
Double-spotted queenfish – Scomberoides lysan – Palau
Dusky gregory – Stegastes nigricans – Guam
Dusky gregory – Stegastes nigricans – Palau
Dusky gregory – Stegastes nigricans – Palau
Eastern triangular butterflyfish – Chaetodon baronessa – Palau
Eclipse butterflyfish – Chaetodon bennetti – Guam
Eclipse butterflyfish – Chaetodon bennetti – Palau
Eight-banded butterflyfish – Chaetodon octofasciatus – Palau
Emperor angelfish – Pomacanthus imperator – Guam
Filament-fin parrotfish – Mini-fin parrotfish – Scarus altipinnis – Guam
Fine-lined surgeonfish – Inshore surgeonfish – Acanthurus grammoptilus – Palau
Fine-spotted rabbitfish – Siganus punctatissimus – Palau
Fire dartfish – Nemateleotris magnifica – Palau
Fivelined cardinalfish – Cheilodipterus quinquelineatus – Palau
Fivestripe wrasse – Thalassoma quinquevittatum – Guam
Flagtail grouper – Cephalopholis urodeta – Palau
Flagtail triggerfish – Sufflamen chrysopterum – Palau
Foursaddle grouper – Epinephelus spilotoceps – Palau
Foxface rabbitfish – Siganus vulpinus – Palau
Freckled hawkfish – Paracirrhites forsteri – Guam
Freckled hawkfish – Paracirrhites forsteri – Palau
Giant moray – Gymnothorax javanicus -Palau
Giant sweetlips – Plectorhinchus albovittatus – Palau
Giant sweetlips – Plectorhinchus albovittatus – Palau
Giant thorny oyster – Spondylus varius – Palau
Giant trevally – Caranx ignobilis – Palau
Goldback damsel – Pomacentrus nigromanus – Palau
Golden damsel – Amblyglyphidodon aureus – Palau
Golden damsel – Amblyglyphidodon aureus – Palau
Golden spadefish – Platax boersii – Palau
Goldsaddle goatfish – Parupeneus cyclostomus – Guam
Goldsaddle goatfish – Parupeneus cyclostomus – Palau
Goldstriped sweetlips – Plectorhinchus chrysotaenia – Palau
Gorham dragonet – Diplogrammus goramensis – Palau
Granulated sea star – Choriaster granulatus – Palau
Gray reef shark – Carcharhinus amblyrhynchos – Palau
Graytail angelfish – Chaetodontoplus poliourus – Palau
Green barrel sea squirt – Didemnum molle – Palau
Hawksbill sea turtle – Eretmochelys imbricata – Palau
Hawksbill sea turtle – Eretmochelys imbricata – Palau
Hinged fangblenny – Crested sabretooth blenny – Petroscirtes mitratus – Palau
Hinged shrimpfish – Jointed razorfish – Aeoliscus strigatus
Honeycomb grouper – Epinephelus merra – Guam
Honeycomb grouper – Epinephelus merra – Palau
Honeycomb grouper – Epinephelus merra – Palau
Humbug dascylus – Dascyllus aruanus – Guam
Humbug dascylus – Dascyllus aruanus – Palau
Humpback snapper – Lutjanus gibbus – Palau
Humpback unicornfish – Naso brachycentron – Palau
Humphead wrasse – Napoleon wrasse – Cheilinus undulatus – Palau
Humphead wrasse – Napoleon wrasse – Cheilinus undulatus – Palau
Humphead wrasse juvenile – Napoleon wrasse – Cheilinus undulatus – Palau
Humpnose bigeye bream – Monotaxis grandoculis – Guam
Humpnose bigeye bream – Monotaxis grandoculis – Guam
Humpnose bigeye bream – Monotaxis grandoculis – Palau
Indian surge damsel – Chrysiptera brownriggii – Guam
Indian surge damsel – Chrysiptera brownriggii – Guam
Inexplicable goby – Cryptocentrus inexplicatus – Palau
Island goatfish – Parupeneus insularis – Guam
Island goatfish – Parupeneus insularis – Palau
Jelly-eating anemone – Entacmaea medusivora – Palau
Jewel damsel – Plectroglyphidodon lacrymatus – Guam
Jewel damsel – Plectroglyphidodon lacrymatus – Palau
Jewelled blenny – Salarias fasciatus – Guam
Jewelled blenny – Salarias fasciatus – Palau
Keyhole angelfish – Centropyge tibicen – Palau
Lagoon triggerfish – Rhinecanthus aculeatus – Guam
Lagoon triggerfish – Rhinecanthus aculeatus – Palau
Latent slingjaw wrasse – Epibulus brevis – Palau
Latent slingjaw wrasse – Epibulus brevis – Palau
Lattice butterflyfish – Chaetodon rafflesi – Palau
Latticed sandperch – Parapercis clathrata – Guam
Lemon damsel – Pomacentrus moluccensis – Palau
Lemonpeel angelfish – Centropyge flavissima – Guam
Lemon sponge – Leucetta chagosensis – Palau
Leopard coral grouper – Leopard coralgrouper – Plectropomus leopardus – Palau
Leopard wrasse – initial phase – Macropharyngodon meleagris – Guam
Leopard wrasse – Macropharyngodon meleagris – Guam
Lined bristletooth – Ctenochaetus striatus – Palau
Lined butterflyfish – Chaetodon lineolatus – Palau
Lined butterflyfish – with odd squiggles – Chaetodon lineolatus – Palau
Lined rabbitfish – Siganus lineatus – Palau
Longfin spadefish – Platax teira – Palau
Long-finned goby – Ocellate glidergoby – Valenciennea longipinnis – Palau
Orange-dashed goby – Valenciennea puella – Palau
Longfin emperor – Lethrinus erythropterus – Palau
Longhorn cowfish – Lactoria cornuta – Guam
Long-jawed mackerel – Longjaw mackel – Rastrelliger kanagurta – Palau
Longnose butterflyfish – Forcipiger flavissimus – Guam
Longnose filefish – Oxymonacanthus longirostris – Guam
Longspot snapper – Lutjanus fulviflamma – Palau
Lowfin chub – Brassy drummer – Kyphosus vaigienses – Palau
Lunar fusilier – Caesio lunaris – Palau
Magenta dottyback – Purple dottyback – Pictichromis porphyrea – Palau
Mandarinfish – Synchiropus splendidus – Palau
Mandarinfish – Synchiropus splendidus – Palau
Mango sponge – Stylissa massa – Palau
Mangrove snapper – Lutjanus argentimaculatus – Guam
Manybar goatfish – Parupeneus multifasciatus – Guam
Manybar goatfish – Parupeneus multifasciatus – Palau
Many-spotted sweetlips – Harlequin sweetlips – Plectorhinchus chaetodonoides – Palau
Many-spotted sweetlips – Harlequin sweetlips – Plectorhinchus chaetodonoides – Palau
Many-spotted sweetlips juvenile – Harlequin sweetlips – Plectorhinchus chaetodonoides – Palau
Marbled cone sea snail – Conus marmoreus – Palau
Masked bannerfish – Heniochus monoceros – Guam
Masked bannerfish – Heniochus monoceros – Palau
Masked grouper – Gracila albomarginata – Palau
Messmate pipefish – Corythoichthys intestinalis – Palau
Messmate pipefish – Corythoichthys intestinalis – Palau
Meyer’s butterflyfish – Chaetodon meyeri – Palau
Midnight snapper juvenile – Macolor macularis – Guam
Midnight snapper – Macolor macularis – Palau
Midnight snapper – Macolor macularis – Palau
Midnight snapper – Macolor macularis – Palau
Mimic surgeonfish – Acanthurus pyroferus – Palau
Monarch damsel – Dischistodus pseudochrysopoecilus – Palau
Monarch damsel juvenile – Dischistodus pseudochrysopoecilus – Palau
Moon jelly – Aurelia aurita – Palau
Moorish idol – Zanclus cornutus – Guam
Moorish idol – Zanclus cornutus – Palau
Mural goby – Mural glidergoby – Valenciennea muralis – Palau
Mural goby – Mural glidergoby – Valenciennea muralis – Palau
Mushroom coral – Fungia fungites – Palau
Neon damsel – Pomacentrus coelestis – Palau
Neon fusilier – Pterocaesio tile – Palau
Onespot damsel – Onespot demoiselle – Chrysiptera unimaculata – Guam
Onestripe wormfish – Blacklined wormfish – Gunnellichthys pleurotaenia – Palau
Orangeband surgeonfish – Acanthurus olivaceus – Guam
Orangefin anemonefish – Amphiprion chrysopterus – Palau
Orangefin anemonefish – Amphiprion chrysopterus – Palau
Orangelined triggerfish – Balistapus undulatus – Guam
Orangelined triggerfish – Balistapus undulatus – Palau
Orangespine unicornfish – Naso lituratus – Guam
Orangespine unicornfish – Naso lituratus – Palau
Orangestriped emperor – Lethrinus obsoletus – Palau
Oriental sweetlips – Plectorhinchus vittatus – Palau
Ornate butterflyfish – Chaetodon ornatissimus- Guam
Ornate butterflyfish – Chaetodon ornatissimus – Palau
Oval-spot butterflyfish – Chaetodon speculum – Palau
Pacific bird wrasse – Gomphosus varius – Guam
Pacific bird wrasse – Gomphosus varius – Palau
Pacific bird wrasse – Gomphosus varius – Palau
Pacific bullethead parrotfish – Chlorurus spilurus – Guam
Pacific bullethead parrotfish – Chlorurus spilurus – Guam
Pacific bullethead parrotfish – Chlorurus spilurus – Guam
Pacific bullethead parrotfish – Chlorurus spilurus – Palau
Pacific bullethead parrotfish – Chlorurus spilurus – Palau
Pacific bullethead parrotfish – Chlorurus spilurus
Pacific bullethead parrotfish juvenile – Chlorurus spilurus – Palau
Pacific double-saddle butterflyfish – Chaetodon ulietensis – Guam
Pacific double-saddle butterflyfish – Chaetodon ulietensis – Palau
Pacific dusky wrasse – Halichoeres annularis – Palau
Pacific dusky wrasse juvenile – Halichoeres annularis – Palau
Pacific longnose parrotfish – Hipposcarus longiceps – Palau
Pacific longnose parrotfish juvenile – Hipposcarus longiceps – Palau
Pacific Ocean crown-of-thorns sea star – Acanthaster solaris – Guam
Pacific Ocean crown-of-thorns sea star – Acanthaster solaris – Palau
Pacific sailfin tang – Zebrasoma veliferum – Guam
Pacific sailfin tang – Zebrasoma veliferum – Palau
Pacific sailfin tang – Zebrasoma veliferum – Palau
Pacific surge damsel – Chrysiptera leucopoma – Guam
Pacific surge damsel – Chrysiptera leucopoma – Palau
Painted spiny lobster – Panulirus versicolor – Palau
Pajama cardinalfish – Sphaeramia nematoptera – Palau
Pajama cardinalfish – Sphaeramia nematoptera – Palau
Pajama cardinalfish – Sphaeramia nematoptera – Palau
Palau golden jelly – Mastigias papua etpisoni – Palau
Palau golden jelly – Mastigias papua etpisoni – Palau
Palenose parrotfish – Scarus psittacus – Guam
Pale-spot damsel – Dischistodus chrysopoecilus – Palau
Pale-spot damsel juvenile – Whitepatch damsel – Dischistodus chrysopoecilus – Palau
Paletail unicornfish – Naso brevirostris – Palau
Pastel-green wrasse – Halichoeres chloropterus – Palau
Pastel-green wrasse – Halichoeres chloropterus – Palau
Pastel-green wrasse – Halichoeres chloropterus – Palau
Pearly monocle bream – Scolopsis margantifera – Palau
Pearly monocle bream – Scolopsis margaritifera – Palau
Peacock grouper – Cephalopholis argus – Palau
Pearl-scale angelfish – Centropyge vroliki – Palau
Pearly monocle bream – Scolopsis margantifera – Palau
Pearly monocle bream – Scolopsis margaritifera – Palau
Pearly monocle bream juvenile – Scolopsis margantifera – Palau
Pennant bannerfish – Heniochus chrysostomus – Guam
Pennant bannerfish – Heniochus chrysostomus – Palau
Pennant bannerfish – Heniochus chrysostomus – Palau
Persian carpet flatworm – Pseudobiceros bedfordi – Palau
Pincushion sea star – Culcita novaeguineae – Palau
Pincushion sea star – Culcita novaeguineae – Palau
Pink anemonefish – Amphiprion perideraion – Palau
Pinktail triggerfish – Melichthys vidua – Guam
Pinktail triggerfish – Melichthys vidua – Guam
Pinktail triggerfish – Melichthys vidua – Palau
Pinstriped wrasse – Halichoeres melanurus – Palau
Pinstriped wrasse – Halichoeres melanurus – Palau
Porcupinefish – Diodon hystrix – Palau
Princesss damsel – Pomacentrus vaiuli – Guam
Pseudoceros imitatus nudibranch – Pseudoceros imitatus – Palau
Purple anthias – Pseudanthias tuka – Palau
Pyramid butterflyfish – Hemitaurichthys polylepis – Palau
Pyramid butterflyfish – Hemitaurichthys polylepis – Palau
Raccoon butterflyfish – Chaetodon lunula – Guam
Red-and-black anemonefish – Amphiprion melanopus – Guam
Red-and-black anemonefish – Amphiprion melanopus – Palau
Redbreasted wrasse – Maori wrasse – Cheilus facsiatus – Guam
Redbreasted wrasse – Cheilus facsiatus – Palau
Red-dotted guard crab – Trapezia cymodoce – Palau
Redfin anthias – Nemanthias dispar – Palau
Redfin anthias – Nemanthias dispar – Palau
Redfin bream subadult – Monotaxis heterodon – Palau
Redfin butterflyfish – Chaetodon lunulatus – Guam
Redfin butterflyfish – Chaetodon lunulatus – Palau
Red lionfish – Pterois volitans – Palau
Redlip parrotfish – Scarus rubroviolaceus – Guam
Redshoulder wrasse – Stethojulis bandanensis – Guam
Redshoulder wrasse – Stethojulis bandanensis – Palau
Red snapper – Lutjanus bohar – Palau
Redtooth triggerfish – Odonus niger – Palau
Reef manta ray – Mobula alfredi – Palau
Regal angelfish – Pygoplites diacanthus – Palau
Reticulated butterflyfish – Chaetodon reticulatus – Guam
Reticulated butterflyfish – Chaetodon reticulatus – Palau
Reticulated dascylus – Dascyllus reticualtus – Palau
Ringtail Maori wrasse – Oxycheilinus unifasciatus – Guam
Ringtail Maori wrasse – Oxycheilinus unifasciatus – Palau
Rockmover wrasse – Novaculichthys taeniourus – Palau
Rockmover wrasse – Novaculichthys taeniourus – Guam
Rockmover wrasse juvenile – Novaculichthys taeniourus – Palau
Roving coral grouper – Plectropomus pessuliferus – Palau
Roving coral grouper – Plectropomus pessuliferus – Palau
Saber squirrelfish – Sargocentron spiniferum – Palau
Saddled butterflyfish – Chaetodon ephippium – Guam
Saddled butterflyfish – Chaetodon ephippium – Palau
Sailfin snapper – Symphorichthys spilurus – Palau
Sailor’s eyeball alga – Bubble alga – Valonia ventricosa – Palau
Sand shrimpgoby – Ctenogobiops feroculus – Palau
Sangi cardinalfish juvenile – Fibramia thermalis – Palau
Scaly damsel – Pomacentrus lepidogenys – Palau
Schlegel’s parrotfish – Scarus schlegeli – Palau
Schroeder’s rainbow wrasse – Coris schroederii – Palau
Schroeder’s rainbow wrasse – Coris schroederii – Palau
Scissortail fusilier – Caesio caerulaurea – Guam
Scissortail sergeant – Abudefduf sexfasciatus – Palau
Scribbled rabbitfish – Siganus spinus – Guam
Sea grapes – Caulerpa racemosa – Palau
Sea krait – Laticauda colubrina – Palau
Segmented blenny – Salarias segmentatus – Palau
Segmented blenny – Salarias segmentatus – Palau
Shadowfin soldierfish – Myripristis adusta – Palau
Shadowfin soldierfish – Myripristis adusta – Palau
Sharksucker – Echeneis naucrates – Palau
Sharksucker – Echeneis naucrates – Palau
Sixbar wrasse – Thalassoma hardwicke – Palau
Slender grouper – Anyperodon leucogrammicus – Palau
Slender lizardfish – Saurida gracilis – Palau
Slingjaw wrasse – Epibulus insidiator – Guam
Slingjaw wrasse – Epibulus insidiator – Guam
Slingjaw wrasse – Epibulus insidiator – Palau
Slingjaw wrasse – Epibulus insidiator – Palau
Slingjaw wrasse – Epibulus insidiator – Palau
Six-banded angelfish – Pomacanthus sextriatus – Palau
Small giant clam – Tridacna maxima – Guam
Smooth giant clam – Tridacna derasa – Palau
Snowflake moray – Echidna nebulosa – Palau
Snowflake moray – Echidna nebulosa – Palau
Snubnosed dart – Trachinotus blochii – Palau
Solander’s toby – Canthigaster solandri – Guam
Spanish flag snapper – Lutjanus carponotatus – Palau
Speckled butterflyfish – Chaetodon citrinellus – Guam
Speckled damsel – Pomacentrus bankanensis – Palau
Spotbanded butterflyfish – Spot-banded butterflyfish – Chaetodon punctatofasciatus – Palau
Spotfin squirrelfish – Neoniphon sammara – Guam
Spotfin squirrelfish – Neoniphon sammara – Palau
Spot-tail butterflyfish – Chaetodon ocellicaudus – Palau
Spotted boxfish – Ostracion meliagris – Palau
Spotted boxfish – Ostracion meliagris – Guam
Spotted coral grouper – Plectropomus maculatus – Palau
Spotted linckia sea star – Linckia multiflora – Palau
Spotted parrotfish – Cetoscarus ocellatus – Palau
Spotted parrotfish initial phase – Cetoscarus ocellatus – Palau
Spotted parrotfish juvenile – Cetoscarus ocellatus – Palau
Squaretail coral grouper – Plectropomus areolatus – Palau
Squaretail mullet – Ellochelon vaigiensis – Palau
Staghorn damsel – Amblyglyphidodon curacao – Guam
Staghorn damsel – Amblygyphidodon curacao – Palau
Starry blenny – Salarias ramosus – Palau
Starry puffer – Arothron stellatus – Guam
Starry puffer – Arothron stellatus – Palau
Starry rabbitfish – Honeycomb rabbitfish – Siganus stellatus – Palau
Steephead parrotfish – Chlorurus microrhinos – Palau
Steephead parrotfish – Chlorurus microrhinos – Palau
Steephead parrotfish – Chlorurus microrhinos – Palau
Striped large-eye bream – Goldspot seabream – Gnathodentex aureolineatus – Guam
Striped large-eye bream – Gnathodentex aureolineatus – Palau
Striped monocle bream – Scolopsis lineata – Guam
Striped puffer – Narrowlined puffer – Arothron manilensis – Guam
Striped surgeonfish – Acanthurus lineatus – Guam
Striped surgeonfish – Acanthurus lineatus – Palau
Sunset wrasse – Thalassoma lutescens – Guam
Surf parrotfish – Scarus rivulatus – Palau
Swarthy parrotfish – Scarus niger – Palau
Synapted sea cucumber – Lion’s paw sea cucumber – Euapta godeffroyi – Guam
Synapted cucumber – Euapta godeffroyi – Palau
Tailspot squirrelfish – Sargocentron caudimaculatum – Palau
Teardrop butterflyfish – Chaetodon unimaculatus – Guam
Teardrop butterflyfish – Chaetodon unimaculatus – Palau
Threadfin butterflyfish – Chaetodon auriga – Guam
Threadfin butterflyfish – Chaetodon auriga – Palau
Threadfin butterflyfish – Chaetodon auriga – Palau
Threadfin butterflyfish – Chaetodon auriga – Palau
Threadfin cardinalfish – Zoramia leptacantha – Palau
Three-lined monocle bream – Scolopsis trilineata – Palau
Three-line wrasse – Stethojulis strigiventer – Palau
Three-spot dascyllus – Threespot dascyllus – Dascyllus trimaculatus – Guam
Threespot wrasse – Halichoeres trimaculatus – Guam
Threespot wrasse – Halichoeres trimaculatus – Guam
Threespot wrasse – Halichoeres trimaculatus – Palau
Threespot wrasse – Halichoeres trimaculatus – Palau
Threespot wrasse – Halichoeres trimaculatus – Palau
Three-striped whiptail – Pentapodus trivittatus – Palau
Three-striped whiptail – Pentapodus trivittatus – Palau
Thrush sneaky wrasse – Pteragogus turdus – Palau
Thumbprint emperor – Lethrinus harak – Palau
Tiger snapping shrimp – Alpheus bellulus – Palau
Tiger snapping shrimp – Alpheus bellulus – Palau
Titan triggerfish – Balistoides viridencens – Palau
Tripletail wrasse – Cheilinus trilobatus – Guam
Tripletail wrasse – Cheilinus trilobatus – Palau
Trumpetfish – Aulostomus chinensis – Palau
Tubelip wrasse – Labrichthys unilineatus – Palau
Two-spined angelfish – Centropyge bispinosa – Palau
Twospot damsel – Chrysiptera biocellata – Palau
Twospot damsel – Chrysiptera biocellata – Palau
Twospot snapper – Lutjanus biguttatus – Palau
Twotone dartfish – Ptereleotris evides – Guam
Twotone dartfish – Ptereleotris evides – Palau
Vagabond butterflyfish – Chaetodon vagabundus – Palau
Variegated lizardfish – Synodus variegatus – Palau
Wedgetail triggerfish – Rhinecanthus rectangulus – Guam
Whitebelly damsel – Amblyglyphidodon leucogaster – Palau
Whitecheek surgeonfish – Acanthurus nigricans – Palau
White damsel – Dischistodus perspicillatus – Palau
White damsel juvenile -Dischistodus perspicillatus – Palau
White-spotted puffer – Arothron hispidus – Palau
White-spotted rabbitfish – Siganus canaliculatus – Palau
White-spotted rabbitfish – Siganus canaliculatus – Palau
White-spotted eagle ray – Aetobatus ocellatus – Palau
Whitespotted grouper – Epinephelus coeruleopunctatus – Palau
Whitetail damsel – Pomacentrus chrysurus – Palau
Whitetail stingray – Urogymnus granulatus – Palau
Whitetip reef shark – Triaenodon obesus – Palau
Wolf cardinalfish – Cheilodipterus artus – Palau
Yaeyama coralblenny – Palespotted combtooth blenny – Ecsenius yaeyamaensis – Palau
Yellow-barred parrotfish – Scarus dimidiatus – Palau
Yellow-barred parrotfish – Scarus dimidiatus – Palau
Yellow-barred parrotfish – Yellowbar parrotfish – Scarus dimidiatus – Palau
Yellow boxfish – Ostracion cubicus – Palau
Yellowfin goatfish – Mulloidichthys vanicolensis – Palau
Yellowlip emperor – Lethrinus xanthochilus – Palau
Yellowmargin triggerfish subadult – Pseudobalises flavimarginatus – Palau
Yellow-mask angelfish – Blueface angelfish – Pomacanthus xanthometopon – Palau
Yellowsaddle goatfish – Goldsaddle goatfish – Parupeneus cyclostomus Palau
Yellowsaddle goatfish – Parupeneus cyclostomus – Palau
Yellowstripe goatfish – Mulloidichthys flavolineatus – Guam
Yellowstripe goatfish – Mulloidichthys flavolineatus – Palau
Yellowtail coris – Coris gaimard – Palau
Yellowtail coris juvenile – Coris gaimard – Palau
Yellowtail damsel – Neoglyphidodon nigroris – Palau
Yellowtail demoiselle – Neopomacentrus azysron – Palau
Yellowtail fusilier – Caesio cuning – Palau
Yellowtail fusilier – Caesio cuning – Palau
Yellowtail tubelip – Diprotacanthus xanthurus – Palau
Yellow tang – Zebrasoma flavescens – Guam
Zebra dartfish – Ptereleotris zebra – Palau
Zebra sea cucumber – Stichopus vastus Palau