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Various species photographed in 2024. All photos copyright by Martin Holladay and Karyn Patno.

Ambon chromis (ambon puller) – Pycnochromis amboinensis (Chromis amboinensis)
Ambon damsel – Pomacentrus amboinensis
Ambon damsel – Pomacentrus amboinensis
Anchor tuskfish – Choerodon anchorago
Arabian cowrie – Mauritia arabica
Argus wrasse – Halichoeres argus
Argus wrasse juvenile – Halichoeres argus
Axilspot wrasse – Halichoeres podostigma
Axilspot wrasse – Halichoeres podostigma
Axilspot wrasse juveniile – Halichoeres podostigma
Azurespot damsel – Pomacentrus grammorhynchus
Azurespot damsel juvenile – Pomacentrus grammorhynchus
Banded boxer shrimp – Stenopus hispidus
Banded goby – Amblygobius phalaena
Banded sea urchin – Echinothrix calamaris
Banded sole – Soleichthys heterorhinos
Barcheek trevally – Carangoides plagiotaenia
Barred thicklip – Hemigymnus fasciatus
Batu coris – Coris batuensis
Bat volute – Cymbiola vespertilio
Bengal sergeant – Abudefduf bengalensis
Bennett’s feather star – Oxycomanthus bennetti
Bicolor angelfish – Centropyge bicolor
Bicolor goatfish – Parupeneus barberinoides
Bigfin reef squid – Sepioteuthis lessoniana
Black-backed butterflyfish – Chaetodon melannotus
Blackbar chromis – Chromis retrofasciata
Blackbar filefish – Pervagor janthinosoma
Blackbar wrasse – Thalassoma nigrofasciatum
Blackbelt hogfish – Bodianus mesothorax
Black damsel – Neoglyphidodon melas
Blackeye thicklip – Hemigymnus melapterus
Black feather sea star – Colobometra perspinosa
Blackheaded filefish – Pervagor melanocephalus
Blacklip butterflyfish – Chaetodon kleinii
Blackpatch triggerfish – Rhinecanthus verrucosus
Black-saddled toby – Canthigaster valentini
Blacksaddle snake eel – Ophichthus cephalozona
Black sea urchin – Diadema setosum
Blackspot shrimpgoby – Cryptocentrus nigrocellatus
Blackspot snapper – Lutjanus ehrenbergii
Black-spotted egg cowrie – Calpumus verrucosus
Blackspotted puffer – Arothron nigropunctatus
Blackspotted puffer – Arothron nigropunctatus
Blackspotted puffer – Arothron nigropunctatus
Blackspotted sea cucumber – Pearsonothuria graeffei
Blackstreak surgeonfish (Eyeline surgeonfish or Eyestripe surgeonfish) – Acanthurus nigricauda
Black-tail sergeant – Abudefduf lorenzi
Blacktip grouper – Epinephelus fasciatus
Blacktip silverbiddy – Gerres oyena
Blackvent damsel – Dischistodus melanotus
Blackvent damsel juvenile – Dischistodus melanotus
Bleeker’s parrotfish – Chlorurus bleekeri
Bleeker’s parrotfish – intermediate phase – Chlorurus bleekeri
Blubberlip snapper – Lutjanus rivulatus
Blueback damsel – Pomacentrus simsiang
Blueback damsel – Pomacentrus simsiang
Blueback damsel juvenile – Pomacentrus simsiang
Blue candy nudibranch (Swollen phyllidia nudibranch) – Phyllidia varicosa
Blue damsel – Pomacentrus pavo
Blue devil female – Chrysiptera cyanea
Blue-green chromis – Chromis viridis
Bluelipped bristletooth – Ctenochaetus cyanocheilus
Bluelipped bristletooth juvenile – Ctenochaetus cyanocheilus
Blue sea star – Linckia laevigata
Bluespine unicornfish – Naso unicornis
Bluespot mullet – Moolgarda seheli
Blue-spotted wrasse – Anampses caeruleopunctatus
Bluestreak cleaner wrasse – Labroides dimidiatus
Bluestreak goby – Valenciennea strigata
Blue swimming crab – Portunus pelagicus
Bluntheaded wrasse (Twotone wrasse) – Thalassoma amblycephalum
Bower’s parrotfish – Chlorurus bowersi
Bracelet cardinalfish – Nectamia viria
Brassy trevally – Caranx papuensis
Bridled monocle bream – Scolopsis bilineata
Bristle-tailed filefish – Acreichthys tomentosus
Broom filefish – Amanses scopas
Brown coral blenny – Atrosalarias fuscus
Brushtail tang – Zebrasoma scopas
Bubble alga (Sailor’s eyeball) – Valonia ventricosa
Bumblebee snail – Engina mendicaria
Burrowing urchin – Echinometra mathaei
Cake urchin – Tripneustes gratilla
Cake urchin – Collector urchin – Tripneustes gratilla
Captain cone sea snail – Conus capitaneus
Chameleon parrotfish – Scarus chameleon
Checkerboard wrasse – Halichoeres hortulanus
Checkerboard wrasse juvenile – Halichoeres hortulanus
Checkered snapper – Lutjanus decussatus
Chocolate chip sea star – Protoreaster nodosus
Chocolate chip sea star – Protoreaster nodosus
Chocolate chip sea star – Protoreaster nodosus
Chocolate chip sea star – Protoreaster nodosus
Christmas tree worm – Spirobranchus giganteus
Cigar wrasse – Cheilio inermis
Circular spadefish – Platax orbicularis
Circle-cheek wrasse – Halichoeres miniatus
Clark’s anemonefish – Amphiprion clarkii
Cockatoo waspfish – Ablabys taenianotus
Cocktail wrasse – Pteragogus flagellifer
Common seahorse – Hippocampus kuda (Hippocampus taeniopterus)
Convict surgeonfish – Acanthurus triostegus
Cornetfish – Fistularia commersonii
Crescent wrasse (Moon wrasse) – Thalassoma lunare
Crocodile needlefish (Houndfish) – Tylosurus crocodilus
Crown-of-thorns sea star – Acanthaster planci
Crown-of-thorns sea star – Acanthaster solaris
Cryptoplax oculatus chiton – Cryptoplax oculatus
Cutribbon wrasse – Stethojulis interrupta
Dash-dot goatfish – Parupeneus barbennus
Day octopus – Octopus cyanea
Day octopus disguising itself as a sea snail shell – Octopus cyanea
Decumbent Spanish dancer – Hexabranchus lacer
Decumbent Spanish dancer eggs – Hexabranchus lacer
Disc coral – Turbinaria mesenterina
Double-bar goatfish – Parupeneus crassilabris
Dusky dottyback – Pseudochromis fuscus
Dusky dottyback – yellow variation – Pseudochromis fuscus
Dusky gregory (Black gregory, Dusky damsel) – Stegastes nigricans
Dwarf hawkfish – Cirrhitichthys falco
Dwarf lionfish – Neochirus brachyptera
Dwarf lionfish – Shortfin lionfish – Neochirus brachyptera
Eastern triangular butterflyfish – Chaetodon baronessa
Episcopal miter sea snail – Mitra mitra
Eight-banded butterflyfish – Chaetodon octofasciatus
Estuarine halfbeak – Zenarchopterus dispar
False clown anemonefish – Amphiprion ocellaris
Feather duster tube worm – Sabellastarte indica
Filamentous blenny – Cirripectes filamentosus
Fine-lined surgeonfish – Acanthurus grammoptilus
Fingerprint toby (Compressed toby) – Canthigaster compressa
Firetail dottyback – Labracinus cyclophthalmus
Fivelined cardinalfish – Cheilodipterus quinquelineatus
Five-lined ribbon worm – Baseodiscus quinquelineatus
Forskals sidegill slug nudibranch – Pleurobranchus forskalii
Fourline wrasse – Stethojulis trilineata
Fourline wrasse – intermediate phase – Stethojulis trilineata
Freckled goatfish – Upeneus tragula
Freckled grouper – Cephalopholis microprion
Freckled hawkfish – Paracirrhites forsteri
Frostfin cardinalfish – Ostorhinchus hoevenii
Galaxy coral – Galaxea fascicularis
Geographic wrasse – Anampses geographicus
Ghost cardinalfish – Nectamia fusca
Giant barrel sponge – Xestospongia testudinaria
Giant carpet anemone – Stichodactyla gigantea
Giant carpet anemone – Stichodactyla gigantea
Golden cuttlefish – Sepia esculenta
Golden damsel – Amblyglyphidodon aureus
Golden rabbitfish – Siganus guttatus
Goniobranchus coi nudibranch – Goniobranchus coi
Gorgeous shrimpgoby – Amblyeleotris wheeleri
Graceful anemone shrimp – Ancylomenes venustus
Greenfin parrotfish (Pacific bullethead parrotfish) – Chlorurus spilurus
Granulated sea star – Choriaster granulatus
Great barracuda – Sphyraena barracuda
Green barrel sea squirt – Didemnum molle
Green brittle star – Ophiarachna incrassata
Green sea turtle – Chelonia mydas
Guineafowl puffer – Arothron meleagris
Halloween hermit crab – Ciliopagurus strigatus
Hammerhead slug (velvet sea slug) – like a nudibranch – Chelidonura varians
Heart urchin – Metalia spatagus
Hebrew cone sea snail – Conus ebraeus
Hinged shrimpfish – Aeoliscus strigatus
Honeycomb coral – Diploastrea heliopora
Honeycomb coral – Diploastrea heliopora
Honeycomb toby (Whitespotted sharpnose puffer) – Canthigaster janthinoptera
Humbug dascyllus – Dascyllus aruanus
Humbug dascyllus – Dascyllus aruanus
Humpback prawn – Metapenaeopsis lamellata
Humphead bannerfish – Heniochus varius
Immaculate puffer – Arothon immaculatus
Indian parrotfish – Chlorurus capistratoides
Indo-Pacific sergeant – Abudefduf vaigiensis
Jansen’s wrasse – Thalassoma jansenii
Jewel damsel – Plectoglyphidodon lacrymatus
Jewelled blenny – Salarias fasciatus
Jewelled blenny – Salarias fasciatus
Johnston damselfish – pale variation – Plectroglyphidodon johnstonianus
Keyhole angelfish – Centropyge tibicen
King damsel – Chrysiptera rex
Lampert’s sea cucumber – Synaptula lamperti
Large tube sponge – Haliclona fascigera
Latent slingjaw wrasse – Epibulus brevis
Latticed butterflyfish – Chaetodon rafflesi
Latticed sandperch – Parapercis clathrata
Lemon damsel – Pomacentrus moluccensis
Lined bristletooth – Ctenochaetus striatus
Lined butterflyfish – Chaetodon lineolatus
Lobster shrimp – Neaxius acanthus
Longarm prawn – Heteropenaeus longimanus
Long-finned goby – Valenciennea longipinnis
Longfin spadefish – Platax teira
Longnose filefish – Oxymonacanthus longirostris
Longspot snapper – Lutjanus fulviflamma
Luzon sea star – Echinaster luzonicus
Magnificent shrimpgoby – Tomiyamichthys emilyae
Mango sponge – Stylissa massa
Manybar goatfish – Parupeneus multifasciatus
Manylined cardinalfish – Ostorhinchuys multilineatus
Many-spotted sweetlips juvenile – Plectorhinchus chaetodonoides
Map puffer – Arothron mappa
Marbled parrotfish – Leptoscarus vaigiensis
Maxima clam – Tridacna maxima
Messmate pipefish – Corythoichthys intestinalis
Milkfish – Chanos chanos
Milkspotted puffer – Chelonodon patoca (Chelonodontops patoca)
Mimic surgeonfish – Acanthurus pyroferus
Mimic surgeonfish juvenile – Acanthurus pyroferus
Monarch damsel – Dischistodus pseudochrysopoecilus
Moorish idol – Zanclus cornutus
Mottled lightfoot crab – Grapsus albolineatus
Mottled lightfoot crab – Grapsus albolineatus
Mottled sea star – Nardoa tuberculata
Mushroom coral – Fungia fungites
Nail sea star – Mithrodia clavigera
Nebulous wrasse (Pink-bellied wrasse, Cloud wrasse) – Halichoeres nebulosus
Nebulous wrasse juvenile – Halichoeres nebulosusi
Neon damsel – Pomacentrus coelestis
Nerita undata sea snail – Nerita undata
Net-like brown algae – Hydroclathrus clathratus
Noble feather star – Comaster nobilis
Noble feather star – Comaster nobilis
Ocellated eagle ray – Aetobatus ocellatus
Onespot damsel – Chrysiptera unimaculata
Onespot damsel juvenile – Chrysiptera unimaculata
Orange anemonefish – Amphiprion sandaracinos
Orange-banded coralfish – Coradion chrysozonus
Orange-lined triggerfish – Balistapus undulatus
Orange socket surgeonfish – Acanthurus auranticavus
Orange socket surgeonfish juvenile – Acanthurus auranticavus
Orangespine unicornfish – Naso lituratus
Orangetip bristletooth – Ctenochaetus tominiensis
Orbicular cardinalfish – Sphaeramia orbicularis
Oval-spot butterflyfish – Chaetodon speculum
Pacific bird wrasse – Gomphosus varius
Pacific sailfin tang – Zebrasoma veliferum
Pale anemone hermit crab – Dardanus deformis
Palenose parrotfish – Scarus psittacus
Pale-spot damsel – Dischistodus chrysopoecilus
Panda butterflyfish – Chaetodon adiergastos
Papuan toby – Canthigaster papua
Papuan toby – Canthigaster papua
Pastel-green wrasse – Halichoeres chloropterus
Pastel-green wrasse – Halichoeres chloropterus
Peacock flounder – Bothus mancus
Peacock grouper – Cephalopholis argus
Peacock-tail anemone shrimp – Periclimenes brevicarpalis
Pearl-scaled angelfish – Centropyge vroliki
Pearly dartfish – Ptereleotris microlepis
Pimpled phyllidiella nudibranch – Phyllidiella pustulosa
Pimpled phyllidiella nudibranch – Phyllidiella pustulosa
Pin-cushion sea star – Culcita novaeguineae
Pin-cushion sea star – Culcita novaeguineae
Pin-cushion sea star – Culcita novaeguineae
Pin-cushion sea star – Culcita novaeguineae
Pin-cushion sea star – Culcita novaeguineae
Pink anemonefish – Amphiprion peridaraion
Pinstriped wrasse – Halichoeres melanurus
Pinstriped wrasse – intermediate phase – Halichoeres melanurus
Pizza anemone – Cryptodendrum adhaesivum
Pizza anemone – Cryptodendrum adhaesivum
Pontifical miter sea snail – Mitra stictica
Porcelain anemone crab – Neopetrolisthes maculatus
Princess damsel – Pomacentrus vaiuli
Purple anthias (male) – Pseudanthias tuka
Pustular triton sea snail – Argobuccinum pustulosum
Quoy’s parrotfish – Scarus quoyi
Raccoon butterflyfish – Chaetodon lunula
Red and black anemonefish – Amphiprion melanopus
Redbreasted wrasse (Maori wrasse) – Cheilus facsiatus
Redfin butterflyfish – Chaetodon lunulatus
Redshoulder wrasse – Stethojulis bandanensis
Redshoulder wrasse – intermediate phase – Stethojulis bandanensis
Red lionfish – Pterois volitans
Reef lizardfish – Synodus varietagus
Regal angelfish – Pygoplites diacanthus
Reticulated dascyllus – Dascyllus reticulatus
Reticulated moray – Richardson’s moray – Gymnothorax richardsonii
Reticulated sandperch – Parapercis tetracantha
Richmond’s wrasse – intermediate phase – Halichoeres richmondi
Ringed pipefish – Dunckerocampus dactyliophorus
Rippled rockskipper – Istiblennius edentulus
Robust silverside – Slender hardyhead – Atherinomorus lacunosus
Rockmover wrasse – Novaculichthys taeniourus
Rockmover wrasse – Novaculichthys taeniourus
Sally lightfoot crab – Mottled lightfoot crab – Grapsus albolineatus
Sand-dusted cone sea snail – Conus arenatus
Scheele’s conger – Tropical conger – Ariosoma scheelei
Scheele’s conger – Tropical conger – Ariosoma scheelei
Scissortail fusilier – Caesio caerulaurea
Scissortail sergeant – Abudefduf sexfasciatus
Scribbled rabbitfish – Siganus spinus
Scribbled rabbitfish – Siganus spinus
Sea krait – Laticauda colubrina
Serpent sea cucumber – Opheodesoma grisea
Sevenstripe cardinalfish – Ostorhinchus novemfasciatus
Sharksucker – Echeneis naucrates
Sharpnose sandperch – Parapercis cylindrica
Sharpnose sandperch – Parapercis cylindrica
Six-banded angelfish juvenile – Pomacanthus sextriatus
Sixbar wrasse – Thalassoma hardwicke
Sixbar wrasse juvenile – Thalassoma hardwicke
Six-lined soapfish – Sixline
soapfish – Grammistes sexlineatus
Sixspot goby – Valenciennea sexguttata
Sixstripe wrasse – Pseudocheilinus hexataenia
Sleepy sponge crab – Dromia dormia
Snake sea cucumber – Synapta maculata
Snake sea cucumber – Synapta maculata
Snowflake moray – Echidna nebulosa
Snowflake moray – Echidna nebulosa
Soldier cone sea snail – Conus miles
Solor boxfish – female – Ostracion solorensis
Solor boxfish – male – Ostracion solorensis
Spaghetti worm – Loimia medusa
Speckled butterflyfish – Chaetodon citrinellus
Speckled damsel – Pomacentrus bankanensis
Speckled damsel – Pomacentrus bankanensis
Speckledfin grouper juvenile – Epinephelus ongus
Spider conch – Lambis lambis
Spinecheek anemonefish – Premnas biaculeatus
Spiny chromis (spiny puller) – Acanthochromis polyacanthus
Spiny devilfish – Inimicus didactylus
Spiny devilfish – Inimicus didactylus
Splitband cardinalfish – Blue-eye cardinalfish – Ostorhinchus compressus
Spotfin lionfish – Pterois antennata
Spotgill cardinalfish – Ostorhinchus chrysopomus
Spot-nape butterflyfish – Chaetodon oxycephalus
Spot-tail butterflyfish – Chaetodon ocellicaudus
Spotted boxfish – Ostracion meliagris
Spotted jelly (Papuan spotted jelly) – Mastigias papua
Spotted sandperch (Brown-blotched sandperch) – Parapercis millepunctata
Springer’s damsel – Chrysiptera springeri
Squaretail mullet – Ellochelon vaigiensis
Squat shrimp (Sexy shrimp) – Thor amboinensis
Staghorn damsel – Amblyglyphidodon curacao
Star puffer – older juvenile – Arothron stellatus
Starry blenny – Salarias ramosus
Striated cone sea snail – Conus striatus
Striped fangblenny – Meiacanthus grammistes
Striped monocle bream – Scolopsis lineata
Striped puffer – Arothron manilensis
Streaky rockskipper – Istiblennius dussumieri
Talbot’s damsel – Chrysiptera talboti
Threadfin anthias – Pseudanthias huchti
Threadfin butterflyfish – Chaetodon auriga
Threeline wrasse – Stethojulis strigiventer
Threeline wrasse juvenile – Stethojulis strigiventer
Threespot damsel juvenile – Pomacentrus tripunctatus
Three-spot dascylus juvenile – Dascyllus trimaculatus
Threespot squirrelfish – Sargocentron cornutum
Thumbprint emperor – Lethrinus harak
Tiger conch sea snail – Conomurex luhuanus
Titan triggerfish – Balistoides viridencens
Tomato anemonefish – Amphiprion frenatus
Tomato anemonefish – Amphiprion frenatus
Toothy cardinalfish – Cheilodipterus isostigmus
Topsail chub – Kyphos cinerascens
Triangular butterflyfish – Chaetodon triangulum
Tricolor parrotfish – Scarus tricolor
Tricolor parrotfish – intermediate phase – Scarus tricolor
Tripletail wrasse – Cheilinus trilobatus
Tripletail wrasse – intermediate phase – Cheilinus trilobatus
Tubelip wrasse – Labrichthys unilineatus
Two-spined angelfish – Centropyge bispinosa
Twotone dartfish – Ptereleotris evides
Twotone dartfish juvenile – Ptereleotris evides
Unomia stolonifera soft coral – Unomia stolonifera
Vagabond butterflyfish – Chaetodon vagabundus
Vanikoro sweeper – Pempheris vanicolensis
Variegated feather duster tube worm – Bispira variegata
Variegated feather duster tube worm – Bispira variegata
Variegated lizardfish – Reef lizardfish – Synodus variegatus
Vermitid worm shell – Ceraesignum maximum
Vexilla vexillum sea snail – Vexilla vexillum
Violet coral shell – Coralliophila violacea
Violet demoiselle – Neopomacentrus violascens
Violetlined parrotfish – Scarus globiceps
Virgate rabbitfish – Siganus virgatus
Warty egg shell cowrie – Calpurnus verrucosus
Warty egg shell cowrie – Calpurnus verrucosus
Wedge sea hare (Blunt end sea hare) – Dolabella auricularia
Weed wrasse – Seagrass wrasse – Halichoeres papilionaceus
Whiteband damsel – Plectroglyphidodon leucozonus
Whiteband damsel juvenile – Plectroglyphidodon leucozonus
Whitebelly toby – Canthigaster bennetti
White-backed shrimpgoby – Cryptocentrus albidorsus
White-backed shrimpgoby – Cryptocentrus albidorsus
White damsel – Dischistodus perspicillatus
White-shouldered whiptail – Pentapodus bifasciatus
White-spotted hermit crab – Dardanus megistos
White-spotted hermit crab – Dardanus megistos
White-spotted puffer – Arothron hispidus
White-spotted puffer – Arothron hispidus
White-spotted rabbitfish – Siganus canaliculatus
Whitestreak monocle bream – Scolopsis ciliata
Whitetail damsel – Pomacentrus chrysurus
Whitetail damsel – Pomacentrus chrysurus
Whitetail damsel juvenile – Pomacentrus chrysurus
Wirenet filefish – Cantherhines pardalis
Yellow-barred parrotfish – Scarus dimidiatus
Yellow-barred parrotfish – intermediate phase – Scarus dimidiatus
Yellow boxfish juvenile – Ostracion cubicus
Yellow chromis – Chromis analis
Yellowfin parrotfish – Scarus flavipectoralis
Yellowfin parrotfish – intermediate phase – Scarus flavipectoralis
Yellowflanked fairy wrasse – Ryukyu fairy wrasse – Cirrhilabrus-ryukyuensis
Yellow-footed hermit crab – Clibanarius virescens
Yellowlined cardinalfish – Ostorhinchus chrysotaenia
Yellowlined cardinalfish – Ostorhinchus chrystaenia
Yellowstriped cardinalfish – Ostorhinchus cyanosoma
Yellowstripe goatfish – Mulloidichthys flavolineatus
Yellowtail coris – Coris gaimard
Yellowtail coris juvenile – Coris gaimard
Yellowtail damsel – Neoglyphidodon nigroris
Yellowtail wrasse (Spotted wrasse) – Anampses meleagrides
Zebra lionfish – Dendrochirus zebra
Zigzag wrasse – Halichoeres scapularis